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The signal transduction mechanisms underlying differentiation and proliferation of porcine preimplantation embryos are not completely understood. In this study, the phosphatidylinositol (PI) 3-kinase/Akt pathway was studied during preimplantation development of the CGR8 endodermal precursor cell line derived from the porcine blastocyst. The effect of the Akt phosphorylation inhibitor, Akt inhibitor VIII (AI VIII), on endodermal cell differentiation was assessed by examination of the expression of endodermal marker proteins. The expression of phosphorylated Akt (p-Akt) was increased in these cells at the time of endoderm differentiation (days 3-7 of development), suggesting a role for the PI 3-kinase/Akt pathway in the regulation of differentiation of these cells. An increase in the expression of phosphorylated glycogen synthase kinase (p-GSK)-3beta was also observed in these cells during differentiation. To assess the role of the PI 3-kinase/Akt pathway in regulating proliferation of endodermal precursor cells, both cell growth and the expression of the proliferation marker, proliferating cell nuclear antigen (PCNA) were examined. A decrease in the expression of p-Akt and p-GSK-3beta was observed at the time of endodermal differentiation
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